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Publication Date

1977

Description

Irrigated and water-stressed swards of perennial ryegrass were compared under field conditions to elucidate the physiological mechanisms which bring about the commonly observed reductions in dry matter (DM) yields caused by drought. Measurements were made during four periods of vegetative growth in the sum­mers of two successive years. The maximum stress developed varied between 62 and 11 6 mm potential soil water deficit. In all but one experimental period, above­ground DM yield was markedly reduced, with corresponding reductions in leaf area index (LAI). The reduction in LAI was caused by a slower rate of leaf expansion> resulting in smal­ler leaves, and a slower rate of appearance. Canopy photosynthesis, on a ground area basis, was markedly lowered by water stress, thus explaining the lower rates ofDM produc­tion. However, the lower rate of photosynthesis could be accounted for by the reduction in leaf area in the stressed swards. Mean rates of photosynthesis, calculated by dividing the rates of canopy photosynthesis by LAI, indi­cated that there was little reduction in the rate of photosynthesis of individual leaves in the stressed compared with irrigated swards. This contrasts with controlled environment studies where a decline in canopy photosynthesis was attributed to a decline in individual leaf photo­synthesis caused by stomata! closure. Calcu­ lation of canopy and leaf resistances from trans­piration measurements confirmed that, in the field, stomata! closure is not a major factor reducing canopy photosynthesis.

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The physiological effects of water stress on perennial ryegrass in the field

Irrigated and water-stressed swards of perennial ryegrass were compared under field conditions to elucidate the physiological mechanisms which bring about the commonly observed reductions in dry matter (DM) yields caused by drought. Measurements were made during four periods of vegetative growth in the sum­mers of two successive years. The maximum stress developed varied between 62 and 11 6 mm potential soil water deficit. In all but one experimental period, above­ground DM yield was markedly reduced, with corresponding reductions in leaf area index (LAI). The reduction in LAI was caused by a slower rate of leaf expansion> resulting in smal­ler leaves, and a slower rate of appearance. Canopy photosynthesis, on a ground area basis, was markedly lowered by water stress, thus explaining the lower rates ofDM produc­tion. However, the lower rate of photosynthesis could be accounted for by the reduction in leaf area in the stressed swards. Mean rates of photosynthesis, calculated by dividing the rates of canopy photosynthesis by LAI, indi­cated that there was little reduction in the rate of photosynthesis of individual leaves in the stressed compared with irrigated swards. This contrasts with controlled environment studies where a decline in canopy photosynthesis was attributed to a decline in individual leaf photo­synthesis caused by stomata! closure. Calcu­ lation of canopy and leaf resistances from trans­piration measurements confirmed that, in the field, stomata! closure is not a major factor reducing canopy photosynthesis.