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Published in PLOS Computational Biology, v. 13, 3, e1005399, p. 1-10.

This is an open access article, free of all copyright, and may be freely reproduced, distributed, transmitted, modified, built upon, or otherwise used by anyone for any lawful purpose. The work is made available under the Creative Commons CC0 public domain dedication.

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The work for this article was supported by the National Science Foundation grant # IIS-1413864.

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