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Published in Acta Neuropathologica, v. 134, issue 5, p. 813-816.

© The Author(s) 2017

This article is distributed under the terms of the Creative Commons Attribution 4.0 International License (, which permits unrestricted use, distribution, and reproduction in any medium, provided you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made.

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This work is supported by the Rainwater Tau Consortium and NIH/NIA (P50AG023501, P01AG019724, T32AG023481-11S1, and P50 AG1657303). Additional support was provided by the Ramón Areces Foundation (C.A.), the Paul G. Allen Family Foundation (A.W.K.), NIH/NIA RC1 AG035610 (G.C.), John Douglas French Alzheimer’s Foundation (G.C.), National Alzheimer’s Coordinating Center Junior Investigator Award (R.S.D.), ASNR Alzheimer’s Imaging Research Grant Program (R.S.D.), RSNA Resident Fellow grant (R.S.D.), and NIH/NIDA grant U24DA041123 (R.S.D.).

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The online version of this article (doi: 10.1007/s00401-017-1764-0) contains supplementary material, which is available to authorized users.

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